When world-famous air racer Dusty Crophopper (voice of Dane Cook) returns to hometown Propwash Junction after another victorious racing season, the former crop duster revels in his new career success until a fateful training run changes his course with a career-ending injury. Forced to shift gears, Dusty decides to train with the Aerial Fire Fighters at Piston Peak Air Attack Base as a Single Engine Air Tanker (SEAT). The heroic, often life-threatening efforts involved are admirable—but seemingly impossible to master—particularly if you’re a crop duster-turned-racer with an injury that can’t be ignored.

Outgoing and spirited super scooper Dipper (voice of Julie Bowen) is skilled at skimming lakes, scooping up more than 1600 gallons of water and dousing angry fires. A former cargo hauler from Alaska, Dipper is an avid air racing fan with a major crush on headline racer Dusty, so she’s head-over-wheels with excitement when the new SEAT shows up at Piston Peak.

Blade Ranger (voice of Ed Harris), a veteran fire-and-rescue helicopter, heads up the Piston Peak Air Attack team. Haunted by a storied past, he’s a tough and demanding air boss with a wry sense of humor, and he’s not exactly enthusiastic about his new trainee Dusty. But Blade is a pro and does everything he can to bring the new SEAT up to speed.

Windlifter (voice of Wes Studi) is a heavy-lift helicopter who can hoist dozens of trees or a huge tank of fire retardant. The American Indian’s wisdom is vast—he’s chock full of fire folklore, and his connection to nature allows him to sense fires before they’re even spotted. The former lumberjack became a firefighter to help others, and no mission is too big for Windlifter.